Adam Tomašových

Earth Science Institute

Slovak Academy of Sciences

Dubravska cesta 9

Bratislava, 84005, Slovakia



Geological Society of America Annual Meeting

European Geoscience Union General Assembly

Ecological Society of America Annual Meeting

Paleontological Society

Paläontologische Gesellschaft

American Society of Naturalists


Data and Sources

Research interests:

(i) Time averaging, stratigraphic mixing and and preservation/fossilization dynamic of skeletal assemblages (southern California shelf, northern Adriatic shelf)

(ii) Macroecology, biogeography, and evolutionary paleobiology of Mesozoic and Cenozoic marine (skeletal) benthic ecosystems at small, regional and provincial scales, mainly along latitudinal, onshore-offshore, and trophic gradients

(iii) Conservation paleobiology/historical ecology of marine benthic communities, using the young fossil record (Holocene)

(iv) Triassic/Jurassic boundary mass extinction – effects of seawater chemistry on extinction, body size and distribution patterns

Current projects:

(i) Tracing changes in seawater temperature during the Late Triassic and the Early Jurassic in the western Tethys supported by the BASE-LiNE Earth ( and the Slovakian VEGA Agency 02/0136/15 [2015-2018]

(ii) Temporal resolution in benthic assemblages: assessing and modeling the roles of burial dynamics, seafloor type, and intrinsic factors using (un)natural experiments, PI – Susan M.Kidwell (University of Chicago) and Clark Alexander (Skidaway Institute of Oceanography), National Science Foundation (NSF EAR 1124318)

Past projects:

(i) The role of articulate brachiopods in Mesozoic benthic communities (Deutsche Forschungsgemeinschaft Grant to Franz T. Fürsich, 2003-2006)

(ii) Global meta-analysis of marine benthic communities (National Science Foundation Grant to Susan Kidwell, 2006-2008)

(iii) Estimating impacts of past natural and anthropogenic disturbances on shelf macrobenthic communities using dead-shell assemblages (University of Southern California Sea Grant Program, 2008-2009) – with Susan Kidwell

(iv) Evaluating and modeling ecological stability and turnover of Jurassic communities of the Tethyan Ocean (Slovak Research and Development Agency (APVV-0248-07) and the Slovakian VEGA Agency [2008-2011]

(v) Bivalves in Time and Space: Testing the accuracy of methods to reconstruct ancestral morphology, dates, geography, and diversification patterns (supported by National Science Foundation)

(vi) Assessing taphonomic and geochemical approaches in evaluating spatial and temporal turnover of marine invertebrate paleocommunitiessupported by Slovak Research and Development Agency (APVV-0644-10) and the Slovakian VEGA Agency [2011-2014]

Mini CV:

Born: 1978

Nationality: Slovak

Comenius University, Department of Geology and Paleontology (Bratislava, Slovakia) – 1996-2002 (MSc)

Würzburg University, Paleontological Institute (Germany) - 2001, 2002 – 2006 (PhD)

University of Chicago, Geophysical Sciences, Research Associate - 11/2006-05/2011

Earth Science Institute, Slovak Academy of Sciences (Slovakia), Researcher - 05-10/2006, 06/2011-present


Tomašových A., Schlögl J., Biroň A., Hudáčková N., and Mikuš T. 2017. Taphonomic clock and bathymetric dependence of cephalopod preservation in bathyal, sediment-starved environments. Palaios, 32, 135-152. PDF.

Tomašových A., Gallmetzer I., Haselmair A., Kaufman D.S., Vidovic J., and Zuschin M. 2017. Stratigraphic unmixing reveals repeated hypoxia events over the past 500 years in the northern Adriatic Sea. Geology 45. PDF. Supplement and data.

Tomašových A., and Jablonski D. 2017. Decoupling of latitudinal gradients in species and genus geographic range size: a signature of clade range expansion. Global Ecology and Biogeography 26, 288-303. Supplement and code-Dryad. PDF.

Bizjack M.T., Kidwell S.M., Velarde R.G., Leonard-Pingel J., Tomašových A. 2017. Detecting, sourcing, and age-dating dredged sediments on the open shelf, southern California, using dead mollusk shells. Marine Pollution Bulletin, 114, 448-465.

Vidović, J., Nawrot, R., Gallmetzer, I., Haselmair, A., Tomašových, A., Stachowitsch, M., Cosovic V. and Zuschin, M. 2016. Anthropogenically induced environmental changes in the northeastern Adriatic Sea in the last 500 years (Panzano Bay, Gulf of Trieste). Biogeosciences, 13, 5965-5981. PDF. Environmental data. Abundance data.

Albano, P.G., Filippova, N., Steger, J., Schmidbaur, H., Tomašových, A., Stachowitsch, M. and Zuschin, M., 2016. Contamination patterns and molluscan and polychaete assemblages in two Persian (Arabian) Gulf oilfields. Marine Ecology-an evolutionary perspective, 37, 907-919.

Tomašových A., Schlögl J., Kaufman D.S., Hudáčková N. 2016. Temporal and bathymetric resolution of nautiloid death assemblages in stratigraphically-condensed oozes (New Caledonia). Terra Nova 28, in press. PDF. Supplement and data.

Tomašových A., Kennedy J.D., Betzner T.J., Bitler Kuehnle N., Edie S., Kim S., Supriya K., White A.E., Rahbek, C., Huang S., Price T.D., and Jablonski D. 2016. Unifying latitudinal gradients in range size and richness across marine and terrestrial systems. Proceedings of the Royal Society B, 20153027. Supplement. PDF.

Tomašových A., Kidwell S.M., and Foygel Barber R. 2016. Inferring skeletal production from time-averaged assemblages: skeletal loss pulls the timing of production pulses towards the modern period. Paleobiology 42, 54-76. PDF. Supplement. Data. Code.

Albano, P. G., Tomašových, A., Stachowitsch, M., and Zuschin, M. 2016. Taxonomic sufficiency in a live–dead agreement study in a tropical setting. Palaeogeography Palaeoclimatology Palaeoecology 449, 341-348.

Albano, P.G., Filippova, N., Steger, J., Kaufman, D.S., Tomašových, A., Stachowitsch, M. and Zuschin, M. 2016. Oil platforms in the Persian (Arabian) Gulf: Living and death assemblages reveal no effects. Continental Shelf Research. Supplement data-composition. Supplement data-age.

Chattopadhyay, D., Zuschin, M. and Tomasovych, A., 2015. How effective are ecological traits against drilling predation? Insights from Recent bivalve assemblages of the northern Red Sea. Palaeogeography, Palaeoclimatology, Palaeoecology 440, 659-670.

Tomašových A., Jablonski D., Berke S.K., Krug Z.A. and Valentine J.W. 2015. Nonlinear thermal gradients shape broad-scale patterns in geographic range size and can reverse Rapoport's rule. Global Ecology and Biogeography 24, 257-267 PDF. Supplement. Code for range expansion. Code for range shuffling.

Tomašových A., Dominici S., Zuschin M. and Merle D. 2014. Onshore–offshore gradient in metacommunity turnover emerges only over macroevolutionary time-scales. Proceedings of the Royal Society B 281, 20141533. Supplement. PDF.

Tomašových A., Kidwell S.M., Foygel Barber R., Kaufman D.S. 2014. Long-term accumulation of carbonate shells reflects a 100-fold drop in loss rate. Geology 42, 9, 819-822. Data Repository with source codes. Raw data. PDF-free-link:

Chattopadhyay D., Zuschin M., and Tomašových A. 2014. Effects of a high-risk environment on edge-drilling behavior: inference from Recent bivalves from the Red Sea. Paleobiology 40, 34-49. PDF-Free.

Vollstaedt H., Eisenhauer A., Wallmann K., Bohm F., Fietzke J., Liebetrau V., Krabbenhift A., Farkas J., Tomašových A., Raddatz J., and Veizer J. 2014. The Phanerozoic δ88/86Sr record of seawater: New constraints on past changes in oceanic carbonate fluxes. Geochimica et Cosmochimica Acta 128, 249-265.

Kidwell S.M. And Tomašových A. 2013. Implications of time-averaged death assemblages for ecology and conservation biology. Annual Reviews of Ecology, Evolution and Systematics 44: 539-563. PDF-Free.

Jablonski, D., Belanger, C.L., Berke, S.K., Huang, S., Krug, A.Z., Roy, K., Tomasovych, A., Valentine, J.W. 2013. Out of the tropics, but how? Fossils, bridge species, and thermal ranges in the dynamics of the marine latitudinal diversity gradient. Proceedings of the National Academy of Sciences of the United States of America 110, 10487-10494.

Berke, S.K., D. Jablonski, A. Z. Krug, K. Roy, A. Tomašových, J. W. Valentine. 2013. Beyond Bergmann’s Rule: size-latitude relationships in marine Bivalvia worldwide. Global Ecology and Biogeography 22, 173-183. PDF-Free.

Šimo V. and Tomašových A. 2013. Trace-fossil assemblages with a new ichnogenus in “spotted” (Fleckenmergel-Fleckenkalk) deposits: a signature of oxygen-limited benthic communities. Geologica Carpathica 64, 355-374.PDF-Free.

Reolid, M., García-García. F., Tomašových, A., Soria, J.M, 2012. Thick brachiopod shell concentrations from prodelta and siliciclastic ramp in a Tortonian Atlantic-Mediterranean Strait (Miocene, Guadix Basin, Southern Spain). Facies 58, 549-571. PDF-Free.

Tomašových A., and Kidwell S.M. 2011. Accounting for the effects of biological variability and temporal autocorrelation in assessing the preservation of species abundance. Paleobiology 37:332-354. PDF-Free

Tomašových A., and Kidwell S.M. 2010. Predicting the effects of increasing temporal scale on species composition, diversity, and rank-abundance distributions. Paleobiology 36:672-695. PDF-Free

Tomašových A., and Kidwell S.M. 2010. The effects of temporal resolution on species turnover and on testing metacommunity models. American Naturalist 175:587-606.PDF-Free

Tomašových A., and Zuschin M. 2009. Variation in brachiopod preservation along a carbonate shelf-basin transect (Red Sea and Gulf of Aden): environmental sensitivity of taphofacies. Palaios 24:697-716. PDF-Free

Tomašových A., and Kidwell S.M. 2009. Fidelity of variation in species composition and diversity partitioning by death assemblages: time averaging transfers diversity from beta to alpha levels. Paleobiology PDF-Free

Tomašových A., and Kidwell S.M. 2009. Preservation of spatial and environmental gradients by death assemblages. Paleobiology PDF-Free

Schlögl, J., Mangold, C., Tomašových A., and Golej, M. 2009. Early and Middle Callovian ammonites from the Pieniny Klippen Belt (Western Carpathians) in hiatal successions: unique biostratigraphic evidence from sediment-filled fissure deposits. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 252:55-79. PDF-Free

Tomašových A., Carlson, S.J., and LaBarbera M. 2008. Ontogenetic niche shift in the brachiopod Terebratalia transversa: relationship between the loss of rotation ability and allometric growth. Palaeontology PDF-Free

Tomašových A. and J. Schlögl. 2008. Analyzing variations in cephalopod abundances in shell concentrations: the combined effects of production and density-dependent cementation rates. Palaios PDF-Free

Tomašových A. 2008: Substrate exploitation and resistance to biotic disturbance in the brachiopod Terebratalia transversa and the bivalve Pododesmus macrochisma. Marine Ecology-Progress Series 363: 157-170. PDF-Free

Tomašových A. 2008: Evaluating neutrality and escalation hypothesis in brachiopod communities from shallow, high-productivity habitats. Evolutionary Ecology Research 10: PDF-Free

Hautmann, M., Benton, M.J., and Tomašových A. 2008. Catastrophic ocean acidification at the Triassic-Jurassic boundary. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 249: 119-127. PDF-Free.

Yanes Y., Tomašových A., Kowalewski M., Castillo C., Aguirre J., Alonso M.R., and Ibáñez M. 2008. Taphonomy and compositional fidelity of Quaternary fossil assemblages of terrestrial gastropods from carbonate-rich environments of the Canary Islands. Lethaia 41:235-256. PDF-Free

Alroy J., and 34 coauthors. 2008. Phanerozoic Trends in the Global Diversity of Marine Invertebrates. Science 5885: 97-100.

Tomašových A. 2008. Composition and distribution of Hettangian brachiopod communities in the West Carpathians: recovery after the end-Triassic mass extinction event. Fossils and Strata 54: 173-181. PDF-Free

van de Schootbrugge, B., Payne, J.L., Tomašových A., Pross, J., Fiebig, J., Benbrahim, M., Föllmi, K.B., Quan, T.M., 2008. Carbon cycle peturbation and stabilization in the wake of the Triassic-Jurassic boundary mass-extinction event. Geochemistry Geophysics Geosystems, v. 9, 10.1029/2007GC001914. Available at AGU - G3 home. PDF-free.

Logan A., Tomašových A., Zuschin M. and Grill B. 2008. Recent brachiopods from the Red Sea and Gulf of Aden. Fossils and Strata 54: 299-309. PDF-Free

Tomašových A. and Siblik M. 2007. Evaluating compositional turnover of brachiopod communities during the end-Triassic mass extinction (Northern Calcareous Alps): removal of dominant groups, recovery and community re-assembly.Palaeogeography, Palaeoclimatology, Palaeoecology 244: 170-200. PDF-Free

Tomašových A. 2006. Brachiopod and bivalve ecology in Late Triassic (Alps, Austria): onshore-offshore replacements caused by variations in sediment and nutrient supply. PALAIOS 21, 344-368 (Paleobiology Database Publication 35). Available at Bioone. PDF-Free

Tomašových A., Fürsich F.T., and Wilmsen M. 2006. Preservation of autochthonous shell beds by positive feedback between increased hardpart-input rates and increased sedimentation rates. Journal of Geology 114: 287-312. Available online via the University of Chicago Press. PDF-Free

Tomašových A. 2006. Linking taphonomy to community-level abundance: insights into compositional fidelity of the Upper Triassic shell concentrations (Eastern Alps). Palaeogeography, Palaeoclimatology, Palaeoecology 235: 355-381. Available at Elsevier. PDF-Free

Tomašových A., Fürsich F. T., and Olszewski T. D. 2006. Modeling shelliness and alteration in shell beds: variation in hardpart-input and burial rates leads to opposing predictions. Paleobiology 32: 278-298.;2 PDF-Free

Tomašových A. 2006. A new Early Jurassic rhynchonellid brachiopod from the western Tethys and implications for systematics of rhynchonellids from the Triassic-Jurassic boundary. Journal of Paleontology 80: 212-228. Available at Bioone. PDF-Free

Tomašových A. and Rothfus T.A. 2005. Differential taphonomy of modern brachiopods (San Juan Islands, Washington State): effect of intrinsic factors on damage and community-level abundance. Lethaia 38: 271-292. PDF-Free

Tomašových A. and Farkaš J. 2005. Cathodoluminescence of Late Triassic terebratulid brachiopods: implications for growth patterns. Palaeogeography, Palaeoclimatology, Palaeoecology 216, 215-233. PDF-Free

Tomašových A. 2004, Postmortem durability and population dynamics affecting the fidelity of size-frequency distributions. PALAIOS 19, 477-496. Bioone. PDF-Free

Tomašových A. 2004, Effect of extrinsic factors on biofabric and brachiopod alteration in a shallow intraplatform carbonate setting (Upper Triassic, West Carpathians). PALAIOS 19, 349-371. Bioone. PDF-Free

Tomašových A. 2004, Microfacies and depositional environment of an Upper Triassic intra-platform carbonate basin: the Fatric Unit of the West Carpathians (Slovakia). Facies 50, 77-105. Availaible at PDF-Free

Kowalewski M., Carroll M., Casazza L., Gupta N., Hannisdal B., Hendy A., Krause R. A. Jr., LaBarbera M., Lazo D.G., Messina C., Puchalski S., Rothfus T. A., Sälgeback J., Stempien J., Terry R.C., Tomašových A. 2003, Quantitative fidelity of brachiopod-mollusk assemblages from modern subtidal environments of San Juan Islands, USA. Journal of Taphonomy 1, 43-65. Available at PDF-Free

Gazdzicki A., Michalík J. and Tomašových A. 2000, Parafavreina coprolites from the Uppermost Triassic of the Western Carpathians. Geologica Carpathica, 51, 245-250. PDF-Free

Short Contributions and NON-ISI publications:

Tomašových A. 2007, Stratigraphic sequences in a storm-dominated, Late Triassic intra-shelf environment of the West Carpathians: implications for correlations with the Eastern Alps. Beringeria 37: 203-216. PDF-Free

A. von Hillebrandt, Krystyn L., Kuerschner W.M., Bown P.R., McRoberts C., Ruhl M., Simms M., Tomasovych A., and Urlichs M. 2007. A candidate GSSP for the base of the Jurassic in the Northern Calcareous Alps (Kuhjoch section, Karwendel Mountains, Tyrol, Austria). International Subcommission on Jurassic Stratigraphy Newsletter 34: 2-20.

Tomašových A. 2000, Lagoonal-peritidal sequences in the Fatra Formation (Rhaetian): an example from the Velká Fatra Mountains (Western Carpathians). Slovak Geological Magazine, 6, 256-259.

Tomašových A. and Michalík J. 2000, Rhaetian/Hettangian passage beds in the carbonate development in the Krížna Nappe (central Western Carpathians, Slovakia). Slovak Geological Magazine, 6, 241-249.

Schlögl J., Aubrecht R. and Tomašových A. 2000, The first find of the Orava Unit in the Púchov section of the Pieniny Klippen Belt (Western Slovakia). Mineralia Slovaca, 32, 45-54.

Tomašových A. 1998, Badenian molluscs from the Devínska Nová Ves - claystone pit (Bratislava, Slovakia). Mineralia Slovaca, 30, 357 - 386.